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Cymatiosphaera hymenosynypha  Morbey  1975 , previously recorded from the Rhaetian Westbury Formation in the United Kingdom, occurs in Triassic–Jurassic boundary strata from Denmark. From the present study it is clear that this small, finely reticulate, ellipsoidal, two-layered palynomorph is not a prasinophyte, but a dinoflagellate cyst with a combination apical/intercalary archaeopyle (type tAtI) which involves three apical plates plus one intercalary plate. This cavate cyst is morphologically similar but not identical to  Valvaeodinium hanneae  Piasecki  2001  from the uppermost Bathonian to lower Callovian of East Greenland, and it is therefore here transferred to  Valvaeodinium , as  Valvaeodinium hymenosynypha  (Morbey) Lindström comb. nov. The appearance of  Valvaeodinium hymenosynypha  in Rhaetian strata in the Danish Basin suggests that the lineage of cavate  Valvaeodinium  cysts originated in the Late Triassic alongside a chorate species of the genus.  Valvaeodinium hymenosynypha  seems to have survived just barely past the end-Triassic mass extinction as it is also present in the earliest Hettangian.

palynology

Valvaeodinium hymenosynypha  (Morbey) comb. nov., a dinoflagellate cyst from the uppermost Triassic and lowermost Jurassic (Rhaetian and Hettangian) of Europe