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Paper electrophoresis and paper chromatography were recently combined under conditions which resulted in a two-dimensional "mapping" procedure for the separation and determination of oligonucleotides.1 This system provides the means to extend previous chemical studies2-4 on mutant strains of tobacco mosaic virus (TMV). The proposed approach is to subject the ribonucleic acid (RNA) of each strain to the action of pancreatic ribonuclease (RNase), and then to separate and analyze the nucleotide and oligonucleotide fragments. Owing to the known specificity of action of RNase, the nature and quantity of the various products obtained by its action provide a clue to the sequential arrangement of nucleotides in the original polynucleotide chains. A similar approach has already been used with strains of TMV,3 4 but the higher resolution of the mapping procedure applied in the present instance allows the comparison of strains to be made in considerably greater detail. We are reporting here the results of such extensive analyses of the RNase digests of the nucleic acids from ordinary TMV and from the M and HR strains.

PRODUCTS OBTAINED BY DIGESTION OF THE NUCLEIC ACIDS OF SOME STRAINS OF TOBACCO MOSAIC VIRUS WITH PANCREATIC RIBONUCLEASE