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Genetic studies showed several years ago that the third chromosomes of wild strains of Drosophila pseudoobscura often carry suppressors of crossing-over. The salivary gland chromosome technique has made it possible to demonstrate not only that these are inverted sections, but also that there are many different inversions present in wild populations inhabiting different geographical regions. So far we have found at least fourteen different gene-sequences in wild stocks, and have found that in most geographical regions several sequences are present, though no single sequence appears to occur throughout the range of the species. There are a number of problems raised by these facts, most of which need further study; the present account is to be regarded only as a preliminary note.

Inversions in the Third Chromosome of Wild Races of Drosophila Pseudoobscura, and Their Use in the Study of the History of the Species