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somes inXXY females.3 Miss Philip added genetic data indicating double crossing-over in the male between the heterochromosomes.4 No combined genetic and cytological data had been presented. Such work will be reported below. During the preparation of the manuscript a preliminary note containing similar evidence was published by Neuhaus.5 We have found cases of "detachment" in males similar to those investigated in females by Kaufmann. The "detachments" represented a separation of an X-chromosome and a small duplication, called Theta (0), which originally were joined together at the fibre point.6 It was possible that the detachment process consisted simply of a breaking-apart of the Xand 0-chromosomes. On the other hand, it seemed probable that the

A Cytogenetic Demonstration of Crossing-Over Between X - and Y - Chromosomes in the Male Drosophila Melanogaster