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The study by Morueta-Holme et al. (1) provides unique and invaluable insight into how tropical montane ecosystems have responded to climate change and other anthropogenic disturbances over the past two-plus centuries. Clearly, one of the most important findings of their study is that many species and ecosystems have shifted their distributions upslope to higher elevations, in accord with rising global mean temperatures (1). Another extremely important finding of the study that is deserving of extra attention is that the high-elevation grasslands of Chimborazo (i.e., the Pajonal) have expanded their distributions not only upslope but also downslope by several hundreds of meters. The upslope …

Downward shift of montane grasslands exemplifies the dual threat of human disturbances to cloud forest biodiversity