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Kline et al. (1) pose a series of “what-if” scenarios intended to cast doubt on our (2) findings. These scenarios arise from a critical misreading of our paper and misinterpretation of the US Department of Agriculture’s Cropland Data Layer (CDL). First, we did not include alfalfa (Medicago sativa), a perennial cultivated-hay legume, in our definition of grassland (2), contrary to Kline et al.’s (1) claim. This error propagates throughout their analysis. Our grassland definition combines all CDL grass-dominated land cover types within the Western Corn Belt (WCB). We agree that grassland increases have occurred in the WCB (3). However, Kline et al. (1) do not address important subregional variability. By our …

Reply to Kline et al.: Cropland data layer provides a valid assessment of recent grassland conversion in the Western Corn Belt