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Smith and Griffin (1) raise four concerns regarding our study of the microbiome of the upper troposphere (2). Sampling microbial cells in this environment remains technically challenging, but the concerns raised are not substantial for the reasons stated below. Our methods were adequate and our conclusions well supported by the available data. In response to Smith and Griffin’s (1) first concern, we provided enough information for our air-sample collection system to be reproduced; the brand of the pump does not represent critical information and can be obtained from the authors upon request. The system has also been described, in detail, previously (3). Briefly, air was collected through a …

Reply to Smith and Griffin: Methods, air flows, and conclusions are robust in the DeLeon-Rodriguez et al. study