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The genetic correlation between two quantitative traits has been assumed to result from the sum of the effects of genes with ++, --, +-, -+, +0, -0,0+, and 0- effects on the two traits in question (see Lerner 1958). Rendel (1963) has suggested a somewhat different approach, namely that the correlation between traits could be considered in terms of the effects of genes on "make" (common resources available to the two traits) and on the distribution of these resources.

The Effects of Different Chromosomes on Four Bristle Number Characters of Drosophila Melanogaster