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Transference of the model-building unit from that of the gene to that of the entire genotype permits the consequences of individual and group selection to be given in terms of parent-offspring covariances. This, in turn, solves two basic problems: (1) extending the genetic analysis to any arbitrary level of complexity; and (2) formulating the prediction equations of genetic advance in terms of parameters which are directly estimable

Selection in Reference to Biological Groups III. Generalized Results of Individual and Group Selection in Terms of Parent-Offspring Covariances