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Species composition, distribution and biomass of benthic macrophyte communities have been studied in a high moun­ tain, oligotrophic, softwater lake from the Central Pyrenees. Isoetids (Isoetes lacustris L., Isoetes selacea Lam., Subitlaria aquatica L. and Eleocharis acicttlaris (L.) Roemer & Schultes) are the dominant macrophytes in areas over 2-3 meters in depth. Natopotamids {Sparganium angtistifolium Michx.) are restricted to shallow waters. Algae (Nitella gracilis (Smith) Agardh) only predominate in deep waters. Three main communities have been distinguished : the Sparganium angustifo­ lium community (160 g dwt nr 2 ), the Isoetes lacustris community (120-460 g dwt nr 2) and the Nitella gracilis community (t 1 g dwt m 2 ). Biomass differences between shallow and deep water Isoetes populations have been attributed to ice-scour stress. Neither irradiance nor slope can explain the lower boundaries observed in the distribution of vascular plants ; sediment features may be responsible for them. Mean lake macrophyte biomass amounts to 140 g dwt m-2 (20 g C nr 2 ), a very high value if compared with boreal lakes with similar limnological characteristics.

Composition, distribution and biomass of benthic macrophyte communities from lake Baciver, a spanish alpine lake in the central Pyrenees