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On the basis of changes in standing crops of live and dead fine roots, we estimated fine-root production on the dry, moderate, and wet sites to be 6.5, 6.3, and 4.8 Mg/ha/year ; turnover to be 7.2, 7.2, and 5.5 Mg/ha/year ; and decomposition to he 8.2, 8.0, and 6.9 Mg/ha/year. The effect of site conditions may be indicated by the number of times that the mean standing crop of live fine roots turned over per year : 2.8 on the dry site, 2.0 on the moderate site, and 1.7 on the wet site. Cyclic death and re- placement of fine roots in a succession of favorable microsites may be an adaptive strategy to maintain the largest number of active roots at a minimum metabolic cost. Results of this study confirm the importance of fine roots as a major pathway of carbon cycling in temperate forests.

Standing crop, production, and turnover of fine roots on dry, moderate, and wet sites of mature Douglas-fir in western Oregon