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Twisted DXCF CBCRs showed dark reversion but relaxed ones did not show detectable dark reversion. The reaction was observed not for the DXCF CBCRs having reversible ligation activity, but for the DXCF CBCRs lacking reversible ligation activity.

A photoproduct of DXCF cyanobacteriochromes without reversible Cys ligation is destabilized by rotating ring twist of the chromophore