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www.nature.com/cr NF-YB1-regulated expression of sucrose transporters in aleurone facilitates sugar loading to rice endosperm Cell Research advance online publication 25 September 2015; doi:10.1038/cr.2015.116 Dear Editor, Cereal endosperms, which consist of an outer aleurone layer and an inner starchy endosperm, provide over 70% staple food for human [www.fao.org]. Rice endosperm development occurs in the caryopsis of a grain. During the grain filling stage, sucrose produced in photosynthetic leaves is transported to grains via the phloem [1, 2]. It has been proposed that sucrose, upon arrival in the caryopsis through the phloem located at the dorsal vascular bundle, is partially hydrolyzed by cell wall invertases (CINs) into glu-cose and fructose [3, 4]. Since endosperm is symplasticall