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The greater horseshoe bat (Rhinolophus ferrumequinum) emitsecholocation sounds consisting of a long constant-frequency (CF) componentpreceeded and followed by a short frequency-modulated (FM) component.When an echo returns with an upward Doppler-shift, the bat compensatesfor the frequency-shift by lowering the emitted frequency in the subsequentorientation sounds and stabilizes the echo image. The bat can accuratelystore frequency-shift information during silent periods of at least severalminutes. The stored frequency-shift information is not affected by tonebursts delivered during silent periods without an overlap with an emittedorientation sound. The system for storage of Doppler-shift information hasproperties similar to a sample and hold circuit with sampling at vocalizationtime and with a rather flat slewing rate for the stored frequency information.

Storage of Doppler-shift information in the echolocation system of the ?CF-FM?-bat,Rhinolophus ferrumequinum