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The whole system of low molecular weight/highly hydrophilic weak-bond linked components of the endosperm of the legume seed of Gleditsia triacanthos was studied on the basis of previous and new data. This reserve incrustant/fibrillar cell wall system may be considered a continuous phase formed by three types of compounds, namely: a) galactomannans/galactomannan-like oligosaccharides highly substituted (reserve) and with low branching (near mannans, fibrillar); b) "mixed-linkage"-glucans (reserve) and cellulose (fibrillar), and c) arabinans (reserve) and low- substituted arabinogalactans (fibrillar). The biological role of these "weak bond-linked" components is not fully understood but they may be multifunctional molecules, as they provide not only the weak bonds necessary for the swelling properties of the endosperm, but also they may contribute to the stabilization of enzymes in the dry endosperm by preventing dehydration.

Weak bond-linked components of the reserve and fibrillar cell walls of the endosperm of Gleditsia triacanthos (Leguminosae)