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The phylogenetic analyses based on molecular data demonstrate that all “micronetine” species of a desmitracheate system form a monophyly.    Macrargus    Dahl, 1886 is a “micronetine” genus, the species of which have a haplotracheate system in general, while    Macrargus   alpinus    Li &amp; Zhu, 1993 was found to have a desmitracheate system; this makes its generic placement problematic. According to the results of phylogenetic analysis, we transfer    M.   alpinus    to another genus as    Nippononeta   alpina    (Li &amp; Zhu, 1993),  comb. n ., and provide a redescription of its genital characters and somatic features. Comparisons with other “micronetine” species with a desmitracheate system are provided. Putative synapomorphies for    Nippononeta   , the clade    Nippononeta    +    Agyneta   , and for the “desmitracheate micronetines” clade, as well as their relationship with    Helophora   , are provided and discussed.

On a desmitracheate “micronetine” Nippononeta alpina (Li & Zhu, 1993), comb. n. ( Araneae , Linyphiidae )