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The small, monophyletic mayfly family   Vietnamellidae   Allen, 1984 has so far only been known from a few extant species of the genus    Vietnamella    Tshernova, 1972, which are all distributed in the Oriental Realm (Vietnam, Thailand, China, and India). Herein we report the first fossil record of   Vietnamellidae   based on a male and female imago from Mid-Cretaceous Burmese amber. We establish the new genus    Burmella    gen. nov.  to accommodate these two new Mesozoic specimens. Their attribution to   Vietnamellidae   is supported by the rounded shape of the hind wings with arched outer margin, the course of thoracic sutures, and characteristics of venation, especially of MP and Cu of the forewings and associated intercalary veins of the cubital field. At the same time,    Burmella    gen. nov.  clearly differs from    Vietnamella    by a diminished number of longitudinal and cross veins in the hind wings, and by the different shape of male genitalia. This first fossil record of   Vietnamellidae   supports an age of at least 100 Ma for this taxon.

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First fossil record of the mayfly family   Vietnamellidae   (  Insecta  ,   Ephemeroptera  ) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution