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Homosexuality occurs in a number of bird species under captive conditions (FISCHER 1965, LILL 1968, RHIJN 1985). However, it appears rare in natural conditions, and is known only in the South African Ostrich Struthio camelus australis (SAUER 1972), Western Gull Larus occidentalis (HUNT & HUNT 1977), Ring-billed Gull L. delawarensis (RYDER et al. 1979, CONOVER et al. 1979, KOVACS & RYDER 1983), California Gull L. californicus (CONOVER et al. 1979), Herring Gull L. argentatus (FITCH 1979), Greylag Goose Anser anser (FABRICIUS 1981), Caspian Tern Sterna caspica (CONOVER 1983), and Common Cormorant Phalacrocorax carbo (FUKUDA unpubl.). Female-female pairings in the gulls are summarized by UEDA (1985). In the course of ethosociological studies of the Black-winged Stilt Himantopus himantopus himantopus (KITAGAWA 1988, in press), three female-female pairs were found during the breeding season, from May to June 1988. Comparison with normal pairs as to nesting territory and incubation activities will be described elsewhere.

Ethosociological studies of the black-winged stilt Himantopus himantopus himantopus. I. Ethogram of the agonistic behaviours.