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This work summarizes the studies about the embryonic development of marsupial frogs that were performed by Eugenia del Pino, an Ecuadorian Biologist, and her collaborators at the Laboratory of Developmental Biology of the Pontifical Catholic University of Ecuador in Quito, with the collaboration of foreign scientists. This analysis summarizes the original contribution of Dr. del Pino to the knowledge of the Developmental Biology of marsupial frogs. Her results were achieved by systematic and well oriented work conducted for more than four decades. For this reason, an epistemological analysis of her research is pertinent and necessary in order to reveal the knowledge construction process in the study of the marsupial frog Gastrotheca riobambae and other frogs in comparison with Xenopus laevis. Eugenia del Pino discovered the features of embryonic development and searched for causal explations. She formulated hypothesis and planned experiments to identify the facts and phenomena in order to formulate a theoretical explanation. All of this was done within a consistent and well supported research line. The present analysis was possible through an exhaustive revision of the publications of Eugenia del Pino, who completed her training as biologist and investigator in universities of the United States of America and Europe. Her contributions have high significance for the development of experimental biology in Ecuador, and have been recognized by the national and international scientific communities. This analysis includes the following themes: Onset of research and outline of the epistemological analysis. The revision includes the reproductive adaptations of the marsupial frog G. riobambae, the modes of oogenesis, and the maternal pouch and its hormonal control. The changes in the maternal pouch for embryonic incubation and the relationship between the embryos and the maternal pouch are reviewed. The embryonic incubation is accompanied by adaptations of the excretory function, changes in the rate of early embryonic development and changes in the process of gastrulation with the formation of an embryonic disk. Body elongation is retarded in marsupial frogs. The current research lines are described. The conclusions highlight the significance of her research.

Esbozo de análisis epistemológico de las investigaciones de Eugenia del Pino sobre el desarrollo animal