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In current investigation we studied the male meiosis and details of secondary   chromosomal associations or pairing recorded for the first time in Dianthus   angulatus Royle ex Benth., from the cold deserts of Lahaul-Spiti (Himachal   Pradesh, India). All the presently studied individuals of the species   existed at 2x level (x = 15). The present chromosome count of n = 15 in the   species is in conformity with the previous counts from India and outside of   India. Secondary associations are defined as the affinity of bivalents to be   positioned in pairs having gentle connections. The secondary chromosomal   associations in the species existed among bivalents/chromosomes were   observed in the meiocytes at metaphase-I and continued till the   metaphase-II. The bivalents positioned side by side and end to end to form   secondary pairing. The difference in the number of bivalents/chromosomes   involved in the secondary associations has also been witnessed. A secondary   association between bivalents is considered to be of immense importance as   it is being taken as a gauge of ploidy in plants. The incidence of such   secondary associations of bivalents/chromosomes in D. angulatus which   existed at 2x level indicated the secondary polyploid nature of the species

Secondary associations in ‘Himalayan pink’ (Dianthus angulatus Royle ex Benth., Caryophyllaceae) from cold deserts of Lahaul-Spiti