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This work was supported by equipment donations from the Russian Foundation for Basic Research (grant 10-04-05084-b) and Alexander von Humboldt Foundation (grant 3.4-8151/07074). A research visit of YM to University of Reading was supported by the Russian Foundation for Basic Research (grant 10-04-01202-а), while the final analysis and publication were supported by Saint Petersburg State University (grant 0.38.292.2015).

Neural crest contribution to the avian shoulder girdle and implications to girdle evolution in vertebrates