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The effects of gri on seed coat and flower color were investigated in a study using Lamprecht line V0400 (PI 527735) as the known source of gri. Seed and flower color data were taken on observations of F 2, BC 1-F2, and BC2,-F2 populations from crosses of V0400 with the recurrent parent S-593. Segregation was observed for a unique flower color pattern: wing petals have a very pale tinge of blue (laelia), and the banner petal has two violet dots (≈3- to 4-mm diameter) on a nearly white background. This very pale laelia flower color cosegregates with gray-white seed coats produced by gri. Furthermore, the very pale laelia color depends on the action of V for expression and is extinguished by v, which produces pure white flowers. Thus, it was demonstrated that the very pale laelia flower color, for which Lamprecht tentatively proposed the gene symbol v pal , is not controlled by an allele at V but is a pleiotropic effect of gri. It

Pleiotropic Effects of gri on Seed Coat and Flower Color in Common Bean