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1 730 597 0.82 110 ± 24 1 920 698 0.76 123 ± 43 1 340 266 0.78 47 ± 11 1z 340 253 0.74 57 ± 12 1z 760 663 0.87 114 ± 37 An important subroutine in any physiological model of the effect of the ambient C O2 concentration (Ca) on plant growth, yield, and water use expresses the effect of Ca on stomatal resistance (Rs). Raschke (1975) postulated that the relevant mechanism is a negative feedback of the internal, intercellular CO2 level (Ci), the increase of which tends to throttle the stomata. Sinclair et al. (1984) proposed that, in effect, this mechanism would maintain a constant ratio of Ci : Ca, equal to ≈0.7 in C3 plants. Earlier, Goudriaan and van Laar (1978) obtained data that suggested the constant ratio hypothesis, as did Wong et al. (1980). Later work by Jones et al. (1985) with soybeans in controlled environment chambers gave ratios between 0.74 and 0.78. In 1984, we did a long-term experiment with greenhouse tomatoes at College Station, Texas, to quantify the effect of CO2 level on growth, water use, and yield (Shaer and van Bavel, 1987). In this experiment, 120 tomato plants (Lycopersicon esculentum, Mill. CV. Tropic) were planted on 25 July 1984 in each of three clear polyethylene chambers, 10 × 2 × 2 m (length/width/ height), that were ventilated at 15 air changes per hour, or 0.0083 m·s-1. The chambers were located inside a polyethylene greenhouse. During the daytime, the level of CO2 was controlled with a CO2 injection system in two chambers, one at 700 and the other at 100 vppm (volume parts of CO2 per million parts of air), with the third one left at ambient level (340 vppm). The plants were grown in 5-liter pots containing 3.5 kg of fritted (calcined) clay, and were kept wellwatered. On 1, 2, and 3 Oct. 1984, simultaneous

Relationships Between Stomatal Resistance and CO2 Level Around and Inside Leaves of Greenhouse Tomatoes