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The brown howler monkey (Alouatta guariba) is a mediumsized and fully arboreal monkey that inhabits the Atlantic Forest of South America. Its geographic distribution extends from southern Bahia through the coastal Brazilian states south to the province of Misiones in northernmost Argentina (Kinzey, 1982; Di Bitetti et al., 1994; Rylands et al., 1988, 1996). Traditionally, two subspecies have been recognized, A. guariba guariba in the north and A. guariba clamitans in the south, although their exact distributions remain unclear. Kinzey (1982) reported that the transition from one subspecies to the other occurs in Espírito Santo and Minas Gerais, in regions flanking the Rio Doce, while Rylands et al. (1988) found evidence suggesting that A. guariba clamitans extends as far north as the Rio Jequitinhonha in northern Minas Gerais, and that A. guariba guariba may be restricted to southern Bahia (see Rylands et al., 1996).

neotropical primates

Cytochrome B Sequences Show Subdivision between Populations of the Brown Howler Monkey (Alouatta guariba) from Rio de Janeiro and Santa Catarina, Brazil