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The multicolored Asian lady beetle Harmonia axyridis (Pallas) is often a pest during autumn, when large numbers take ßight and subsequently land on and enter buildings. Open Þeld experiments were conducted during the autumn ßight to examine the role of linear contrast in visually attracting beetles. White targets printed with 61 by 15-cm stripes that varied in orientation and degree of contrast were covered in insect adhesive, attached to a white background panel, and erected in sites known to be attractive to ßying beetles. Flights occurred when temperatures rose above 21C. Diurnally, peak ßight occurred between 1400 and 1600 hours but shifted progressively earlier within that framework as the ßight season advanced. Beetles signiÞcantly chose targets with high contrast, black stripes; the number of beetles landing on all other contrast levels did not differ signiÞcantly from controls. Vertically positioned stripes attracted more beetles than horizontal ones, but not signiÞ- cantly. The effects of high contrast were absolute rather than relative. The number of beetles landing on targets with 50% contrast stripes did not increase when these were the highest contrast targets available. Results suggest that visual intensity contrast is key to understanding the behavior of H. axyridis in autumn and call into question some commonly accepted ideas regarding choice of overwintering sites. First, during migratory ßight,H.axyridis is not visually attracted to the color white per se, and second, it is unlikely that, during ßight, pheromones are involved in beetle orientation.

Role of Visual Contrast in the Alighting Behavior ofHarmonia axyridis(Coleoptera: Coccinellidae) at Overwintering Sites