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In the butterfly Inachis io, a pupal melanization reducing factor (PMRF) which is located throughout the entire central nervous system controls the intensity of pigmentation of pupal cuticle depending on the background color of the pupation site. PMRF does not only reduce melanization but, in addition, enhances lutein incorporation in a dose-dependent manner to form pupae with yellow color on bright backgrounds.

Involvement of Cyclic Adenosine Monophosphate in the Regulation of Pupal Color Adaptation of the Butterfly, Inachis io