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Ecosystems that possess physical, chemical and biological elements interacting in ways necessary for sustainability are said to have integrity. While conceptually appealing, measuring the condition of ecosystems has proven difficult. Ecosystems are thought to respond to stressors (e.g. detrimental or disorganizing influences) through . changes in functional attributes such as energy flow and nutrient cycling and through changes in community structure as well as general system-level changes (Margalef 1981; Odum 1985; Kay and Schneider 1993). Attempts to assess ecosystem condition have rarely considered energy flow and focused instead on either community structure or nutrient cycling (Karr 1993).

Assessing Ecosystem Integrity Through Energy Flow in Wetlands of Grand Teton and Yellowstone National Parks