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By considering subexponential contributions in large deviation theory, wedetermine the fine structure in the probability distribution of the observabledisplacement of a bead coupled to a molecular motor. More generally, for anystochastic motion along a periodic substrate, this approach reveals a discretesymmetry of this distribution for which hidden degrees of freedom lead to aperiodic modulation of the slope typically associated with the fluctuationtheorem. Contrary to previous interpretations of experimental data, the meanforce exerted by a molecular motor is unrelated to the long-time asymptotics ofthis slope and must rather be extracted from its short-time limit.

Fine-structured large deviations and the fluctuation theorem: Molecular motors and beyond