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l Dr Hadwen Trust for Humane Research, 84a Tilehouse Street, Hitchin, Hertfordshire SG5 2DY, UK; 2Psychology Institute, Life and Health Sciences, Aston University, Aston Triangle, Birmingham, West Midlands B4 7ET, UK; 3Royal Society for the Prevention of Cruelty to Animals, The Causeway, Horsham, West Sussex RH12 lHG, UK; <Rheumatic Diseases Centre, Clinical Sciences Building, Hope Hospital, University ofManchester, Eccles Old Road, Salford, M68HD, UK; 5The Open University, Walton Hall, Milton Keynes, Buckinghamshire MK7 6AA, UK; 6Section of Gastrointestinal Science, Clinical Sciences Building, Hope Hospital, University ofManchester, Eccles Old Road, Salford M6 8HD, UK; 7NMR Research Unit, Institute ofNeurology, University College London, Queen Square, London WC1N 3BG, UK; 8Department of Experimental Psychology, University of Oxford, South Parks Road, Oxford OXl 3UD, UK

Volunteer Studies Replacing Animal Experiments in Brain Research