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Fairly strong positive relationships between stiffness and density have often been reported. No stronger relationships have been found when using parameters of density profiles based on an earlywood-latewood boundary. In this study, we attempt to model the relationships among the stiffness of different samples and simple parameters derived from microdensity profiles, not established according to an earlywood-latewood boundary. Furthermore, we try to determine if there is a genetic variation for the relationship between stiffness and density. From the results, we find that the strongest relationship between a single density parameter and stiff- ness is r0.78, whereas it is r = 0.37 when involving a classical within-ring density parameter. At clone level, r 2 ranges from 0.88 to 0.95, while it is 0.51 for the bulked samples. The mathematical form of the models differ from one clone to another: there is a

annales des sciences forestières

Genetic control of stiffness of standing Douglas fir; from the standing stem to the standardised wood sample, relationships between modulus of elasticity and wood density parameters. Part II